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Owls exhibit specialized hearing functions and ear shapes that also aid in hunting. They are noted for asymmetrical ear placements on the skull in some genera.

Owls can have either internal or external ears, both of which are asymmetrical. Asymmetry has not been reported to extend to the middle or internal ear of the owl.

Asymmetrical ear placement on the skull allows the owl to pinpoint the location of its prey. This is especially true for strictly nocturnal species such as the barn owls Tyto or Tengmalm's owl.

This time difference between ears is about 30 microseconds. Behind the ear openings are modified, dense feathers, densely packed to form a facial ruff, which creates an anterior-facing, concave wall that cups the sound into the ear structure.

The facial disk also acts to direct sound into the ears, and a downward-facing, sharply triangular beak minimizes sound reflection away from the face.

The shape of the facial disk is adjustable at will to focus sounds more effectively. The prominences above a great horned owl's head are commonly mistaken as its ears.

This is not the case; they are merely feather tufts. The ears are on the sides of the head in the usual location in two different locations as described above.

While the auditory and visual capabilities of the owl allow it to locate and pursue its prey, the talons and beak of the owl do the final work.

The owl kills its prey using these talons to crush the skull and knead the body. The Tasmanian masked owl has some of the proportionally longest talons of any bird of prey; they appear enormous in comparison to the body when fully extended to grasp prey.

The family Tytonidae has inner and central toes of about equal length, while the family Strigidae has an inner toe that is distinctly shorter than the central one.

The beak of the owl is short, curved, and downward-facing, and typically hooked at the tip for gripping and tearing its prey. Once prey is captured, the scissor motion of the top and lower bill is used to tear the tissue and kill.

The sharp lower edge of the upper bill works in coordination with the sharp upper edge of the lower bill to deliver this motion.

The downward-facing beak allows the owl's field of vision to be clear, as well as directing sound into the ears without deflecting sound waves away from the face.

The coloration of the owl's plumage plays a key role in its ability to sit still and blend into the environment, making it nearly invisible to prey.

Owls tend to mimic the coloration and sometimes the texture patterns of their surroundings, the barn owl being an exception.

The snowy owl Bubo scandiacus appears nearly bleach-white in color with a few flecks of black, mimicking their snowy surroundings perfectly, while the speckled brown plumage of the tawny owl Strix aluco allows it to lie in wait among the deciduous woodland it prefers for its habitat.

Likewise, the mottled wood-owl Strix ocellata displays shades of brown, tan and black, making the owl nearly invisible in the surrounding trees, especially from behind.

Usually, the only tell-tale sign of a perched owl is its vocalizations or its vividly colored eyes. Most owls are nocturnal , actively hunting their prey in darkness.

Several types of owls, however, are crepuscular —active during the twilight hours of dawn and dusk; one example is the pygmy owl Glaucidium. A few owls are active during the day, also; examples are the burrowing owl Speotyto cunicularia and the short-eared owl Asio flammeus.

Much of the owls' hunting strategy depends on stealth and surprise. Owls have at least two adaptations that aid them in achieving stealth.

First, the dull coloration of their feathers can render them almost invisible under certain conditions. Secondly, serrated edges on the leading edge of owls' remiges muffle an owl's wing beats, allowing an owl's flight to be practically silent.

Some fish-eating owls, for which silence has no evolutionary advantage, lack this adaptation. An owl's sharp beak and powerful talons allow it to kill its prey before swallowing it whole if it is not too big.

Scientists studying the diets of owls are helped by their habit of regurgitating the indigestible parts of their prey such as bones, scales, and fur in the form of pellets.

These "owl pellets" are plentiful and easy to interpret, and are often sold by companies to schools for dissection by students as a lesson in biology and ecology.

Owl eggs typically have a white colour and an almost spherical shape, and range in number from a few to a dozen, depending on species and the particular season; for most, three or four is the more common number.

In at least one species, female owls do not mate with the same male for a lifetime. Female burrowing owls commonly travel and find other mates, while the male stays in his territory and mates with other females.

The systematic placement of owls is disputed. For example, the Sibley—Ahlquist taxonomy of birds finds that, based on DNA-DNA hybridization , owls are more closely related to the nightjars and their allies Caprimulgiformes than to the diurnal predators in the order Falconiformes ; consequently, the Caprimulgiformes are placed in the Strigiformes, and the owls in general become a family, the Strigidae.

A recent study indicates that the drastic rearrangement of the genome of the accipitrids may have obscured any close relationship of theirs with groups such as the owls.

Some to extant species of owls are known, subdivided into two families: 1. Some entirely extinct families have also been erected based on fossil remains; these differ much from modern owls in being less specialized or specialized in a very different way such as the terrestrial Sophiornithidae.

The Paleocene genera Berruornis and Ogygoptynx show that owls were already present as a distinct lineage some 60—57 million years ago Mya , hence, possibly also some 5 million years earlier, at the extinction of the non-avian dinosaurs.

This makes them one of the oldest known groups of non- Galloanserae landbirds. The supposed " Cretaceous owls" Bradycneme and Heptasteornis are apparently non- avialan maniraptors.

During the Paleogene , the Strigiformes radiated into ecological niches now mostly filled by other groups of birds. By the early Neogene , the other lineages had been displaced by other bird orders, leaving only barn-owls and typical owls.

The latter at that time were usually a fairly generic type of probably earless owls similar to today's North American spotted owl or the European tawny owl ; the diversity in size and ecology found in typical owls today developed only subsequently.

Around the Paleogene-Neogene boundary some 25 Mya , barn-owls were the dominant group of owls in southern Europe and adjacent Asia at least; the distribution of fossil and present-day owl lineages indicates that their decline is contemporary with the evolution of the different major lineages of true owls, which for the most part seems to have taken place in Eurasia.

In the Americas, rather, an expansion of immigrant lineages of ancestral typical owls occurred. The supposed fossil herons "Ardea" perplexa Middle Miocene of Sansan, France and "Ardea" lignitum Late Pliocene of Germany were more probably owls; the latter was apparently close to the modern genus Bubo.

Judging from this, the Late Miocene remains from France described as "Ardea" aureliensis should also be restudied. The taxa often united under Strigogyps [34] were formerly placed in part with the owls, specifically the Sophiornithidae; they appear to be Ameghinornithidae instead.

For fossil species and paleosubspecies of extant taxa , see the genus and species articles. For a full list of extant and recently extinct owls, see the article List of owl species.

Among the Kikuyu of Kenya , it was believed that owls were harbingers of death. If one saw an owl or heard its hoot, someone was going to die.

In general, owls are viewed as harbingers of bad luck, ill health, or death. The belief is widespread even today. In Mongolia, the owl is regarded as a benign omen.

In one story, Genghis Khan was hiding from enemies in a small coppice when an owl roosted in the tree above him, which caused his pursuers to think no man could be hidden there.

In modern Japan, owls are regarded as lucky and are carried in the form of a talisman or charm. The modern West generally associates owls with wisdom and vigilance.

This link goes back at least as far as Ancient Greece , where Athens , noted for art and scholarship, and Athena , Athens' patron goddess and the goddess of wisdom, had the owl as a symbol.

Thiselton-Dyer , in his Folk-lore of Shakespeare , says that "from the earliest period it has been considered a bird of ill-omen," and Pliny tells us how, on one occasion, even Rome itself underwent a lustration , because one of them strayed into the Capitol.

He represents it also as a funereal bird, a monster of the night, the very abomination of human kind. Virgil describes its death-howl from the top of the temple by night, a circumstance introduced as a precursor of Dido 's death.

Ovid , too, constantly speaks of this bird's presence as an evil omen; and indeed the same notions respecting it may be found among the writings of most of the ancient poets.

In Hinduism , an owl is the vahana mount of the goddess Lakshmi. People often allude to the reputation of owls as bearers of supernatural danger when they tell misbehaving children, "the owls will get you", [49] and in most Native American folklore, owls are a symbol of death.

For example:. Encouraging natural predators to control rodent population is a natural form of pest control, along with excluding food sources for rodents.

Placing a nest box for owls on a property can help control rodent populations one family of hungry barn owls can consume more than 3, rodents in a nesting season while maintaining the naturally balanced food chain.

Although humans and owls frequently live together in harmony, there have been incidents when owls have attacked humans. All owls are listed in Appendix II of the international CITES treaty the Convention on Illegal Trade in Endangered Species of Wild Fauna and Flora.

Although owls have long been hunted, a news story from Malaysia indicates that the magnitude of owl poaching may be on the rise.

In November , TRAFFIC reported the seizure of plucked and "oven-ready" owls in Peninsular Malaysia. Said Chris Shepherd, Senior Programme Officer for TRAFFIC's Southeast Asia office, "This is the first time we know of where 'ready-prepared' owls have been seized in Malaysia, and it may mark the start of a new trend in wild meat from the region.

We will be monitoring developments closely. Included in the seizure were dead and plucked barn owls, spotted wood owls, crested serpent eagles, barred eagles, and brown wood owls, as well as 7, live lizards.

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